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Brown Algae

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A stipe is a stalk or stemlike structure present in an alga. It may grow as a short structure near the base of the alga (as in Laminaria), or it may develop into a large, complex structure running throughout the algal body (as in Sargassum or Macrocystis). In the most structurally differentiated brown algae (such as Fucus), the tissues within the stipe are divided into three distinct layers or regions. These regions include a central pith, a surrounding cortex, and an outer epidermis, each of which has an analog in the stem of a vascular plant. In some brown algae, the pith region includes a core of elongated cells that resemble the phloem of vascular plants both in structure and function. In others (such as Nereocystis), the center of the stipe is hollow and filled with gas that serves to keep that part of the alga buoyant. The stipe may be relatively flexible and elastic in species like Macrocystis pyrifera that grow in strong currents, or may be more rigid in species like Postelsia palmaeformis that are exposed to the atmosphere at low tide.

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