Tanis reveals that Markus was the first vampire, one of the three sons of Alexander Corvinus, the first immortal. Markus was bitten by a bat and metamorphosed into a vampire, while William was bitten by a wolf and metamorphosed into a werewolf. The third son remained human and gave rise to a line of mortal descendants including Michael, who became the first Lycan–Vampire hybrid. The first werewolves created by William were entirely animal and unable to return to their human forms. Due to William's destructiveness, Markus approached Viktor, then a dying mortal warlord, and offered to metamorphose him and his army into vampires in exchange for tracking down and stopping William, and in destroying those he had infected. Viktor did not kill the brothers because he was led by Markus to believe that doing so would result in the immediate extinction of all other vampires and his Lycan slaves. Tanis further reveals that Selene's father was the architect who built William's prison and that the pendant is the key. Viktor killed Selene's family because they knew the prison's location, but metamorphosed Selene into a vampiress with the location of the prison encoded in her blood. Tanis then refers Selene and Michael to Macaro for help. After Selene and Michael leave, Markus arrives and drinks Tanis' blood to learn Selene and Michael's location, killing Tanis.